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                    Scarabaeoidea of Southern South America  Links   | 
                   
                  
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                  Scarab Guide Links  
                    (goes to the University of Nebraska 
                    web site)   | 
                 
                
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                    Classification  | 
                 
                
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              . 
  Aulacopalpus viridis Guérin-Méneville, 1838  
              Aulacopalpus viridis Guérin-Méneville 1838a:58.  | 
             
            
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                    Figure 18. Parameres in caudal view of 
                        Aulacopalpus viridis.  | 
                   
                  
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                    Distribution of Aulacopalpus pygidialis, Aulacopalpus valdiviensis, 
                    and Aulacopalpus viridis.  | 
                   
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              Description. Male (n=78): Length 11.7-16.3 mm. Width 6.5-8.9 mm. Color: dorsally
                light to dark olive green, sometimes greenish-brown; venter brownish-yellow to 
                yellowish-brown. Head: Figure 5. Dorsal surface densely punctate, sparsely setose, punctures
                moderately large; setae long, slender, tawny to cream-colored. Frontoclypeal suture 
                complete, weakly bisinuate. Labrum moderately produced medially with triangular tooth.
                Maxillary palpus with terminal segment greatly enlarged, elongate (larger than antennal 
                club), with deep sulcus. Mentum with surface moderately setose, apex reflexed into oral
                cavity. Antenna 9- or 10-segmented (segments 5 and 6 sometimes fused), club slightly 
                shorter that combined basal segments. Pronotum: midline with apical half weakly depressed.
                Surface moderately to densely punctate with moderately-sized punctures; moderately 
                setose in apical third and along lateral edges; setae long, slender, tawny to creamcolored.
                Scutellum: Shape parabolic: 1.3 times wider than long medially. Surface glabrous, 
                moderately punctate with moderately-sized punctures. Elytron: Surface glabrous,
                longitudinal striae punctate, moderately impressed; punctures moderately-sized, separated 
                by 2-6 puncture widths; interval sparsely punctate with moderately-sized punctures.
                Epipleuron with row of dense setae just ventral to bead medially, absent in apical and 
                basal 1/5. Pygidium: Width 2.1 times length medially. Surface sparsely punctate, sparsely
                to moderately setose; punctures small; setae cream-colored. Venter: Thorax densely setose; 
                setae long, cream-colored. Legs: Protibia with 3 subequally large teeth in apical
                half. Tarsal claws with modified claw slightly thickened when compared with other claw,
                apex bifurcate. Tarsomere 5 without ventromedial tooth. Meso- and metatibia with apical spurs slender, acute. Mesotibial apex with 12-
                13 spinules. Metatibial apex with 17-20 spinules. Parameres: Figure 18. Basally with
                broad, medial furrow. 
                 
                Female (n=20): Length 13.3-15.3 mm. Width 7.1-9.0 mm. As male except in the
                following respects. Head: Maxillary palpus with terminal segment not enlarged or elongate 
                (smaller than antennal club). Legs: Tarsal claws with modified claw with ventral
                tooth, not thickened when compared with other claw, apex not bifurcate. Meso- and metatibia with apical spurs broader, blunter. | 
             
            
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              Diagnosis. This species is distinguished from all other species in the genus
                Aulacopalpus by the following combination of characters: antennal club not
                elongated (length much shorter than head); male maxillary palpus with terminal
                segment greatly elongated (length greater than antennal club length); 
                head, pronotum, and elytron light to dark olive green; all male tarsal claws
                with modified claw slightly thickened when compared with other claw and
                with apex bifurcate; all female tarsal claws with modified claw with ventral
                tooth and not thickened when compared with other claw; and tarsomere 5 
                without ventromedial tooth; parameres basally without broad, medial furrow. | 
             
            
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              Distribution (see map). Santiago to Malleco, Chile. 
                 
                  Locality Data. Based on 98 specimens examined from AMNH, CASC,
                CMNC, CNCI, FMNH, MGFT, MNNC, UMRM, USNM. 
                 
                CHILE (98). ARAUCO (23): San Alfonso (above Caramávida). LINARES
                (26): Linares. MALLECO (27): Angol, Collipulli, Mininco. MAULE (7): Tregualemu.
                ÑUBLE (1): Chillán. SANTIAGO (7): El Canelo, Leyda. VALDIVIA
                (1): Valdivia. NO DATA (2). | 
             
            
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              Temporal Data. April (2), May (2), June (1), August (2), September (24),
October (24) November (3), December (13). | 
             
            
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              Remarks. The males of A. viridis are easily separated from other species
                of Aulacopalpus due to the greatly enlarged terminal segment of the maxillary
                palpus. Two aspects of the original description by Guérin-Méneville (1838)
                and redescriptions by Guérin-Méneville (1844) and Burmeister (1844) led to
                confusion over the true identity of this species: 1) it was described as having
                "simple, unequal tarsal claws," which is incorrect (the tarsal claws are split
                in males and toothed in females); 2) the type locality for the species was listed
                as Lima, Peru, which is also incorrect (the specimen[s] examined by Guérin-Méneville were probably collected in central Chile). Solier (1851) and Blanchard
                (1851) both remarked that A. viridis was actually from Chile noting that 
                "entomologists frequently confuse insect collections originating from the two
                regions (Peru and Chile) even though they are so different." The confusion
                probably led Philippi and Philippi to describe Tribostethes virens (a synonym
                of A. viridis) in 1864. Tribostethes virens was described using one female
                specimen that has been lost or destroyed since it is not in the Museo Nacional
                de Historia Natural, Santiago, Chile. Philippi and Philippi (1864) described the
                original type for T. virens as a female with a mainly greenish dorsal surface
                and "maxillary palpus very large with the apical segment almost 1.5 times as 
                long as the other segments, with a deep furrow extending almost the whole
                length of the segment." Ohaus (1905) commented, "I determined that Tribostethes
                virens F. Philippi is synonymous with Aulacopalpus viridis; the description 
                of this species fits exactly with fresh specimens of the latter species."
                After careful examination of the original description and many specimens, I
                agree with Ohaus’s decision to synonymize T. virens with A. viridis, and I
                selected an appropriate neotype for both names. | 
             
            
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              Literature cited. 
                Blanchard, C. E. 1851. Ordre des Coleoptera [pp. 129–240]. In: Muséum d'Histoire
                  Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes.
                  vol. 1, part 2 (H. Milne-Edwards, C. E. Blanchard, and H. Lucus, editors). Gide
                  and Baudry, Paris. 
                   
                Burmeister, H. 1844. Handbuch der Entomologie, vol. 4, part 1. T. C. F. Enslin, Berlin.
                  586 pp. 
                   
                Guérin-Méneville, F. E. 1838. Insectes du voyage de la Favorite. Magasin de Zoologie
                8(9):1–80 
                 
                Guérin-Méneville, F. E. 1844. Iconographie du Règne Animal de G. Cuvier, ou Représentation d'après Nature de L'une des Espèces les plus Remarquables et souvent
                non encore Figurees, de Chaque Genre d’Animaux, vol. 3. J. B. Baillière, Paris.
                576 pp. 
                 
                Ohaus, F. 1905. Revision der amerikanischen Anoplognathiden (Coleoptera lamellicornia).
                Stettiner Entomologische Zeitung 66:120–167. 
                 
                Philippi, R. A., and F. Philippi. 1864. Beschreibung einiger neuen Chilenischen Käfer.
                Stettiner Entomologische Zeitung 25:313–406. 
                 
                Solier, A. J. J. 1851. Orden III. Coleopteros [pp. 5–285]. In: Historia Fisica y Politica
                de Chile. Zoología, vol. 5 (C. Gay, editor). C. Gay, Paris. 564 pp. | 
             
            
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              Excerpt from: 
                Smith, 
          A. B. T. 2002. Revision of the South American Endemic genus Aulacopalpus Guérin-Ménville with phylogenetic and biogeographic 
          analyses of the subtribe Brachyisternina (Coleoptera: Ruteliinae: 
          Anoplognathini: Brachysternina). Coleopterists Bulletin 56: 379-437. | 
             
            
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