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                    Scarabaeoidea of Southern South America  Links   | 
                   
                  
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                  Scarab Guide Links  
                    (goes to the University of Nebraska 
                    web site)   | 
                 
                
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                    Classification  | 
                 
                
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              . 
Lichnia gallardoi Gutiérrez, 1943 
              Lichnia gallardoi Gutirrez 1943:151 (valid name). 
              Lichnia porteri Gutirrez 1943:152.  | 
             
            
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              Fig. 9a. Caudal (i) and lateral (ii) views of parameres  
                of Lichnia gallardoi.  | 
             
            
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                    Map of central Chile showing distribution of  
                      Lichnia gallardoi and L. limbata.                        | 
                     
                  
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              Description. 
                Male. Length 5.6-7.6 mm; width 3.5-4.7 mm. Color: Head, pronotum, venter, and 
                legs black; elytral disc testaceous, margin dark brown to black. Head: Surface punctate; 
                punctures small, dense, setose; setae long, moderately dense to dense, white. Clypeus with 
                surface flat, punctate; punctures small, dense, setose; setae short, half as long as clypeal 
                setae, dense, white or black; apex truncate. Labrum with surface punctate; punctures 
                small, dense, setose; setae short, half as long as clypeal setae, dense, white or black; apex 
                rounded with minute, recurved tooth at middle. Mandible on external surface with 
                moderately long, dense, black setae. Labial margins with moderately long, dense, black 
                setae. Eye canthus setose; setae long, dense, black. Antenna with pedicel setose, elongate, 
                segment 2 globular, 3-6 subequal in length, 7-9 lamellate; club small, subequal in length 
                to segments 3-6. Pronotum: Surface punctate; punctures small, moderately 
                dense, with long, moderately dense to dense, white setae. Anterior margin setose; setae 
                short, dense, yellow. Lateral margins slightly rounded, setose; setae long, dense, white. 
                Posterior margin setose; setae long, dense, white. Scutellum: Surface setose; setae long, 
                dense, white. Elytron: Surface depressed posteriorly, punctate; punctures small, 
                moderately dense medially, dense laterally; surface with both stout, black and slender, 
                white setae. Propygidium: Surface exposed, covered with long, dense, white setae; setae 
                overlaying basal 1/4 of pygidium, apices forming straight row. Pygidium: Surface slightly 
                convex, setose; setae long, dense, black. Venter: Thorax punctate; punctures small, dense, 
                with both black and white setae. Mes- and metepisternum setose; setae long, dense, white. 
                Abdominal segments densely setose on lateral margin, sparsely setose medially; setae 
                long, white. Terminal abdominal segment setose on apical half; setae long, dense, black. 
                Margin of apex sinuate. Legs: Surface setose; setae long, moderately dense, both black and 
                white. Protibia slender, with 3 outer teeth and 1 inner apical tooth. Parameres:
                Fig. 9a. 
                 
                Female. Females differ from males in the following respects: length 5.7-7.4 mm; width 
                3.6-4.9 mm. Color: Head, pronotum, venter, and legs reddish brown to black; elytral disc 
                testaceous to red, margin dark brown to black. Head: Clypeal apex weakly rounded, 
                without apical tooth. Pronotum: Lateral margin broadly rounded. Elytron: Surface not 
                depressed posteriorly. Propygidium: Surface setose; setae yellow to yellowish brown, 
                covering 1/3 of pygidium, not dense and not forming a straight row. Pygidium: Apex attenuate. Venter: Apex of terminal sternite of abdomen deeply emarginate. Legs: Protibia 
                expanded. | 
             
            
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              Diagnosis. 
                Lichnia gallardoi resembles Lichnia limbata. Males of L. gallardoi are easily 
                diagnosed by a shorter antennal club  and the presence of a clypeal tooth (males of L. limbata have a longer antennal club and lack a clypeal tooth). The clypeal 
                tooth may be worn down and not visible in many specimens. When using male antennal 
                characters to distinguish L. gallardoi from L. limbata, it must first be determined that the 
                specimens are males by comparing the protibia, which is slender in males and
                expanded along the entire length in females. Otherwise, males of L. gallardoi can 
                be mistaken for females of L. limbata because both have short antennae. On 
                average, both sexes of L. gallardoi are slightly larger than those of L. limbata. With the 
                exception of size, females of these two species are indistinguishable and must be identified 
                based on association with males and geographic location. The distribution of L. limbata is 
                Region V (Valparaíso) to Region VII (Maule), whereas L. gallardoi is found further north 
                in Region IV (Coquimbo) (see map). | 
             
            
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              Remarks. 
                Gutiérrez (1943) differentiated L. porteri from L. gallardoi based on characters such 
                as color of setae and bristles and size difference. After studying a larger series (248 
                specimens) than Gutiérrez had access to, I conclude that these characters represent 
                variation within a single species. Following Article 24.2 of the Code (International 
                Commission on Zoological Nomenclature 1999), I, as first revisor, designate L. gallardoi as the valid name for the species because this name is most commonly used in collections 
                and it better promotes stability. Consequently, L. porteri is here designated as a junior 
                synonym of L. gallardoi. | 
             
            
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              Distribution. 
                Central Chile in Region IV (Coquimbo) (see map). | 
             
            
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              Locality data. 
                248 specimens were examined from ABTS, CASC, CMNC, CNCI, EMUS, FMNH, 
                JEBC, JMEC, LACM, MNHN, MNNC, UCCC, UMRM, USNM, VMDC. 
                 
                CHILE. REGION IV (COQUIMBO) (199): Canela Baja (4), Condoriaco (2), Costa 
                Las Cardas (8), El Molle (4), El Tofo (1), Guanaqueros (1), Huentelauquén (2), Hurtado 
                (2), Illapel (48), La Serena (2), Limari (4), Los Hornitos (27), Los Mantos (1), Los Molles 
                (1), Mincha (21), Ovalle (2), Parque Nacional Fray Jorge (30), Puerto Oscuro (5), 
                Quebrada El Arrayan (2), Rio Limari (5), San Lorenzo (4), Socos (1), Tongoy (6), Vicuna 
                (1), no data (15). NO DATA (49).  | 
             
            
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              Temporal data. 
                January (7), August (2), September (16), October (125), November (43), December
                (6). | 
             
            
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              Excerpt from: 
                  Hawkins, S. J. 2006. A revision of the Chilean tribe Lichniini Burmeister, 1844 
                  (Coleoptera: Scarabaeidae: Melolonthinae). Zootoaxa 1266:1-63.  | 
             
            
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