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              Scarabaeoidea of Southern South America  Links  
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                  Scarab Guide Links  
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                    Classification  | 
                 
                
                  
                    
                      Superfamily... 
                        Family........... 
                        Subfamily .....  
                        Tribe............ 
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                      Scarabaeoidea 
                        Lucanidae 
                        Aesalinae 
                      Ceratognathini | 
                     
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              Hilophyllus martinezi Paulsen & Mondaca, 2006  
              Hilophyllus martinezi Paulsen & Mondaca, 2006: 16.   | 
             
            
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                    Hilophyllus martinezi Paulsen & Mondaca.  | 
                   
                  
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                    Figure 21. Ventral view of male genitalia, H. martinezi.  | 
                   
                  
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                    Figure 25. Localities from which Hilophyllus specimens were examined: H. penai (asterisks), H.
                      argentinensis (squares), and H. martinezi (triangle).  | 
                   
                  
                  
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              Description. Holotype male. Length: 8.0 mm. Width: 2.9 mm. Color/Surface: Dark reddishbrown,
                weakly shining. Dorsum almost uniformly covered with large punctures, 
                most with a single elongate, apically acute scale. Head: Eyes large, width of
                eyes together in dorsal view greater than narrowest interocular width. Mandibles small,
                externally rounded, not abruptly curved inward at middle; dorsal tooth acute; ventral tooth
                of right mandible minute (apex of right mandible broken). Antennal club segments
                elongate, longer than 1.5 times length of head. Pronotum: Sides strongly 
                explanate, explanate margin subequal to eye in width. Anterior angle produced beyond
                anterior margin. Lateral margin broadly rounded, widest before middle. Elytra:
                Shape elongate. Almost uniformly punctate, most punctures with scales, punctures
                without scales forming longitudinal rows (not patches), scales brown and testaceous.
                Sutural interval and lateral margin elevated apically. Surface between punctures 
                roughened, appearing wrinkled at 40x. Legs: Mesotibiae lacking external tooth.
                Male genitalia: See Fig. 21. Parameres with 2 small, external teeth at apex,
                median lobe cylindrical toward apex, not laterally compressed.  
                 
                Female unknown. | 
             
            
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              Diagnosis. 
                This species is distinguished from H. penai by its unarmed mesotibiae. It is, however,
                externally quite similar to H. argentinensis. The most notable external difference between
                these two species is in the anterior angles of the pronotum, which are produced beyond the
                anterior pronotal margin in H. martinezi but more or less contiguous with the anterior
                margin of the pronotum in H. argentinensis. This gives the overall impression of
                a smaller pronotum in H. martinezi. Ideally we would prefer more material upon which to
                base a new species. However, the form of the male genitalia  in H.
                argentinensis and H. martinezi are vastly different. Based on the constancy of male
                genitalic form (i.e., lack of variation) observed in H. penai (n = 4), as well as in H.
                argentinensis (n = 9) across its range, we believe that the unique male genitalic form of H.
                martinezi is compelling evidence that this is a distinct species. | 
             
            
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              Distribution.                CHILE. The species is known from central Chile. | 
             
            
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              | Locality data: 1 specimen examined.
                CHILE (1). VIII Región (1): Los Lleuques.
                Temporal distribution. November (1). | 
             
            
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              Etymology. 
                The species is named martinezi in honor of Antonio Martínez and his contributions to
                the knowledge of South American Coleoptera. | 
             
            
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              Remarks. 
                This single specimen from Los Lleuques, Ñuble, is similar externally to H.
                argentinensis. However, the male genitalia are distinct. The apex of each
                paramere bears two minute, external teeth  rather than the large external tooth 
                present on H. argentinensis. Furthermore, the parameres are strongly narrowed before the
                apex and the median lobe is broad and subcylindrical to the apex, not strongly laterally 
                compressed. Nothing is known about the biology of this species. | 
             
            
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              Excerpt from: 
                Paulsen, M. J. & J. Mondaca E. 2006. Revision of the South American Ceratognathini (Coleoptera: 
                Lucanidae: Aesalinae) with the description of a new genus and 
                a new species. Zootaxa 1191: 1–19.  | 
             
            
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