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                    Scarabaeoidea of Southern South America  Links  
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                    Scarab Guide Links  
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                      Classification  | 
                   
                  
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Leiopsammodius Rakovic, 1981 
Leiopsammodius Rakovic, 1981: 16.  | 
             
            
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                    Leiopsammodius indefensus (Schmidt) 
                      Photo by Paul Skelley.  | 
                   
                  
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              Type species: Psammodius laevicollis Klug, 1845 by original designation. | 
             
            
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              Diagnosis: Body usually robust. Head weakly to distinctly granulate. Clypeus not dentate. 
                Eyes normally developed, not reduced. Elytral intervals margined at base. Metafemur robust. Metatibia 
                dilated apically. Pronotum with a reduced number of ridges, usually two.  | 
             
            
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              Remarks: Leiopsammodius is found nearly worldwide and has about 40 species. Rakovic (1990) 
                reviewed the New World taxa, and most species were also discussed by Cartwright (1955) as Psammodius. 
                Adults of few species are frequently collected at lights, but their habits are unknown. All species appear to be 
                associated with sandy soils. | 
             
            
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              Species: (click on species name for information) 
                Leiopsammodius indefensus (Schmidt) 
                      Leiopsammodius placidus (Schmidt)                  | 
             
            
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              References: 
                Cartwright, O.L. (1955) Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States 
                National Museum, 104(3344), 413–462. 
                 
                Rakovic, M. (1990) Review of the genus Leiopsammodius Rakovic on the world basis with a key to species from the Western Hemisphere 
                and description of a new species (Coleoptera, Scarabaeidae, Aphodiinae). Annotationes Zoologicae et Botanicae, 197, 1–18. | 
             
            
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              Excerpt from: 
                Smith, A. B. T. and P. E. Skelley. 2007. A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa 1458: 1-80. | 
             
            
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