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                    Scarabaeoidea of Southern South America  Links  
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                    Scarab Guide Links  
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                      Classification  | 
                   
                  
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              . Aphodius Illiger, 1798 
                      Aphodius Illiger, 1798: 15  | 
             
            
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                        Aphodius (Calamosternus) granarius (Linnaeus). Male.  
                          Photo by Paul Skelley.  | 
                        Aphodius (Calamosternus) granarius (Linnaeus). Female.  
                          Photo by Paul Skelley. | 
                       
                      
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              Type species: Scarabaeus fimetarius Linnaeus, 1758 by subsequent designation (Latreille 1810). | 
             
            
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              Remarks: The genus Aphodius is a large and taxonomically difficult group that is currently in the process 
            of being re-classified into dozens of genera. We found two species representing two subgenera of Aphodius as 
            occurring in southern South America. Both of these species are introduced to southern South America and 
            native to Europe and Africa. | 
             
            
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              Subgenus Aphodius (Calamosternus) Motschulsky, 1859 
                Calamosternus Motschulsky, 1859: 156. 
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                Type species: Scarabaeus granarius Linnaeus, 1767 by monotypy. 
                Synonyms: Megalisus Mulsant and Rey, 1870: 209; type species: Aphodius stercorarius Mulsant and 
            Rey, 1870 by subsequent designation (Dellacasa 1988). Pseudesymus d’Orbigny, 1896: 218; type species: Aphodius lucidus Klug, 1845 by monotypy. 
            Diagnosis: Color uniformly black. Body robust. Head trituberculate. Pentagonal scutellum 
            usually depressed below the surface of the elytra. Metatibia with apical spurs separated by metatarsus. 
             
            Remarks: Full description and illustrations of Calamosternus and its type species is present in Dellacasa 
            et  
            al. (2001). 
            . 
            Species: (click on species name for information)  
            Aphodius (Calamosternus) granarius (Linnaeus) | 
             
            
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              Subgenus Aphodius (Labarrus) Mulsant and Rey, 1870 
                Aphodius (Labarrus) Mulsant and Rey, 1870: 516. 
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                Type species: Scarabaeus lividus Olivier, 1789 by monotypy. 
                Synonym: Aphodius (Pseudocalamosternus) Balthasar, 1936: 1; type species: Aphodius (Pseudocalamosternus) rigidus Balthasar, 1936 by monotypy. 
                Diagnosis: Body robust. Pale brown and black color pattern is similar only to members of Aidophus in
            South America. Head with tubercles on frontal suture. Pronotum lacking basal 
marginal line. Scutellum pentagonal. Metatibia with apical spurs separated by metatarsus. 
Remarks: A full description and illustrations of Labarrus and its type species are present in Dellacasa et 
            al. (2001). Members of the genus Aidophus in South America have similar color pattern to Aphodius (Labarrus) 
            species. Aidophus are easily distinguished by lacking the frontal tubercles, having a triangular 
            scutellum, and in having the metatibial spurs adjacent. 
            . 
            Species: (click on species name for information)  
            Aphodius (Labarrus) pseudolividus Balthasar | 
             
            
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              References: 
                      Dellacasa, M. (1988) Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae 
                (Coleoptera Scarabaeoidea). Memorie della Societa Entomologica Italiana, 66, 1–455. [Dated 1987]. 
                Dellacasa, G., Bordat P. & Dellacasa, M. (2001) A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera: Aphodiidae). 
                  Memorie della Societa Entomologica Italiana, 79, 1–482. 
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              Excerpt from: 
                Smith, A. B. T. and P. E. Skelley. 2007. A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa 1458: 1-80.  | 
             
            
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